The Permian period, 280 to 230 million years ago, was named for the Perm Province of the Ural Mountains in Russia. The Permian signaled the end of the "ancient life" Paleozoic era.

In the Permian, the close ties between geology and evolution were especially apparent. The two great land masses of the Paleozoic drifted close enough together to form one supercontinent, Pangaea. Collisions in the tectonic plates created extensive volcanic activity and heaved up the Urals, Alps, Appalachians, and Rocky Mountains. The shallow inland seas drained to leave deposits of gypsum and salt. Vast sand dunes throughout much of what is now North America and Europe were recorded by massive yellow sandstones (hardened sand dunes) that contained few fossils other than scorpions.

Great glaciers scoured the southern regions of Africa, India, and Australia, further inhibiting life. Conifers and a few cold-hardy plants grew along the fringes of the immense ice...