1889-1981

American physician, anatomist, endocrinologist, and embryologist whose research into the female endocrine system proved the hormonal basis of menstruation. After receiving his M.D. from Johns Hopkins University in 1913, he taught anatomy at the University California at Berkeley, Johns Hopkins, and the University of Rochester before becoming Director of Embryology at the Carnegie Institution of Washington, D.C., in 1940. He and Willard Myron Allen reported their joint discovery of progesterone in "Physiology of the Corpus Luteum" (1929).