Who is the main character in The Red Queen: Sex and the Evolution of Human Nature by Matt Ridley?

There isn't a main character in a narrative sense but The Red Queen is an allusion to a character in Alice In Wonderland. In the story, Alice comes upon a queen running along who implores Alice to run fast. Despite how fast Alice runs, she stays in the same place however because the landscape moves at the same pace that she is. The metaphor is meant to suggest a kind of treadmill where no matter how fast one runs, they stay in the same place.