Insectivorous Plants eBook

This eBook from the Gutenberg Project consists of approximately 527 pages of information about Insectivorous Plants.

Insectivorous Plants eBook

This eBook from the Gutenberg Project consists of approximately 527 pages of information about Insectivorous Plants.

The valve slopes into the cavity of the bladder, or upwards in fig. 18.  It is attached on all sides to the bladder, excepting by its posterior margin, or the lower one in fig. 19, which is free, and forms one side of the slit-like orifice leading into the bladder.  This margin is sharp, thin, and smooth, and rests on the edge of a rim or collar, which dips deeply into the [page 400] bladder, as shown in the longitudinal section (fig. 20) of the collar and valve; it is also shown at c, in fig. 18.  The edge of the valve can thus open only inwards.  As both the valve and collar dip into the bladder, a hollow or depression is here formed, at the base of which lies the slit-like orifice.

The valve is colourless, highly transparent, flexible and elastic.  It is convex in a transverse direction, but has been drawn (fig. 19) in a flattened state, by which its apparent breadth is increased.  It is formed,

Fig. 20. (Utricularia neglecta.) Longitudinal vertical section through the ventral portion of a bladder; showing valve and collar. v, valve; the whole projection above c forms the collar; b, bifid processes; s, ventral surface of bladder.

according to Cohn, of two layers of small cells, which are continuous with the two layers of larger cells forming the walls of the bladder, of which it is evidently a prolongation.  Two pairs of transparent pointed bristles, about as long as the valve itself, arise from near the free posterior margin (fig. 18), and point obliquely outwards in the direction of the antennae.  There are also on the surface of the valve numerous glands, as I will call them; for they have the power of absorption, though I doubt whether they ever secrete.  They consist of three kinds, which [page 401] to a certain extent graduate into one another.  Those situated round the anterior margin of the valve (upper margin in fig. 19) are very numerous and crowded together; they consist of an oblong head on a long pedicel.  The pedicel itself is formed of an elongated cell, surmounted by a short one.  The glands towards the free posterior margin are much larger, few in number, and almost spherical, having short footstalks; the head is formed by the confluence of two cells, the lower one answering to the short upper cell of the pedicel of the oblong glands.  The glands of the third kind have transversely elongated heads, and are seated on very short footstalks; so that they stand parallel and close to the surface of the valve; they may be called the two-armed glands.  The cells forming all these glands contain a nucleus, and are lined by a thin layer of more or less granular protoplasm, the primordial utricle of Mohl.  They are filled with fluid, which must hold much matter in solution, judging from the quantity coagulated after they have been long immersed in alcohol or ether.  The depression in which the valve lies is also lined with innumerable glands; those at the sides having oblong heads and elongated pedicels, exactly like the glands on the adjoining parts of the valve.

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Insectivorous Plants from Project Gutenberg. Public domain.