interval of time, only a slight effect. On the
other hand, the leaves which had been immersed in the
solutions of the chloride of rubidium and magnesium,
of acetate of strontium, nitrate of barium, and citric
acid, were quickly acted on by the phosphate.
Now was water absorbed from these five weak solutions,
and yet, owing to the presence of the salts, did not
prevent the subsequent action of the phosphate?
Or [page 216] may we not suppose* that the interstices
of the walls of the glands were blocked up with the
molecules of these five substances, so that they were
rendered impermeable to water; for had water entered,
we know from the ten trials that the phosphate would
not afterwards have produced any effect? It further
appears that the molecules of the carbonate of ammonia
can quickly pass into glands which, from having been
immersed for 20 m. in a weak solution of sugar, either
absorb the phosphate very slowly or are acted on by
it very slowly. On the other hand, glands, however
they may have been treated, seem easily to permit the
subsequent entrance of the molecules of carbonate of
ammonia. Thus leaves which had been immersed
in a solution (of one part to 437 of water) of nitrate
of potassium for 48 hrs.—of sulphate of
potassium for 24 hrs.—and of the chloride
of potassium for 25 hrs.—on being placed
in a solution of one part of carbonate of ammonia to
218 of water, had their glands immediately blackened,
and after 1 hr. their tentacles somewhat inflected,
and the protoplasm aggregated. But it would
be an endless task to endeavour to ascertain the wonderfully
diversified effects of various solutions on Drosera.
Alcohol (one part to seven of water).—It
has already been shown that half-minims of this strength
placed on the discs of leaves do not cause any inflection;
and that when two days afterwards the leaves were given
bits of meat, they became strongly inflected.
Four leaves were immersed in this mixture, and two
of them after 30 m. were brushed with a camel-hair
brush, like the leaves in the solution of camphor,
but this produced no effect.
* See Dr. M. Traube’s curious experiments on
the production of artificial cells, and on their permeability
to various salts, described in his papers: “Experimente
zur Theorie der Zellenbildung und Endosmose,”
Breslau, 1866; and “Experimente zur physicalischen
Erklrung der Bildung der Zellhaut, ihres Wachsthums
durch Intussusception,” Breslau, 1874.
These researches perhaps explain my results. Dr.
Traube commonly employed as a membrane the precipitate
formed when tannic acid comes into contact with a
solution of gelatine. By allowing a precipitation
of sulphate of barium to take place at the same time,
the membrane becomes “infiltrated” with
this salt; and in consequence of the intercalation
of molecules of sulphate of barium among those of the
gelatine precipitate, the molecular interstices in
the membrane are made smaller. In this altered
condition, the membrane no longer allows the passage
through it of either sulphate of ammonia or nitrate
of barium, though it retains its permeability for
water and chloride of ammonia. [page 217]