nor moves, so that nature was forced to make these
communications for the nutrition of the lungs.
This is plainly false; for simple inspection of the
incubated egg, and of embryos just taken out of the
uterus, shows that the heart moves in them precisely
as in adults, and that nature feels no such necessity.
I have myself repeatedly seen these motions, and Aristotle
is likewise witness of their reality. “The
pulse,” he observes, “inheres in the very
constitution of the heart, and appears from the beginning
as is learned both from the dissection of living animals
and the formation of the chick in the egg.”
[Footnote: Lib de Spiritu, cap. v.] But we further
observe that the passages in question are not only
pervious up to the period of birth in man, as well
as in other animals, as anatomists in general have
described them, but for several months subsequently,
in some indeed for several years, not to say for the
whole course of life; as, for example, in the goose,
snipe, and various birds and many of the smaller animals.
And this circumstance it was, perhaps, that imposed
upon Botallus, who thought he had discovered a new
passage for the blood from the vena cava into the
left ventricle of the heart; and I own that when I
met with the same arrangement in one of the larger
members of the mouse family, in the adult state, I
was myself at first led to something of a like conclusion.
From this it will be understood that in the human
embryo, and in the embryos of animals in which the
communications are not closed, the same thing happens,
namely, that the heart by its motion propels the blood
by obvious and open passages from the vena cava into
the aorta through the cavities of both the ventricles,
the right one receiving the blood from the auricle,
and propelling it by the pulmonary artery and its continuation,
named the ductus arteriosus, into the aorta; the left,
in like manner, charged by the contraction of its
auricle, which has received its supply through the
foramen ovale from the vena cava, contracting, and
projecting the blood through the root of the aorta
into the trunk of that vessel.
In embryos, consequently, whilst the lungs are yet
in a state of inaction, performing no function, subject
to no motion any more than if they had not been present,
nature uses the two ventricles of the heart as if
they formed but one, for the transmission of the blood.
The condition of the embryos of those animals which
have lungs, whilst these organs are yet in abeyance
and not employed, is the same as that of those animals
which have no lungs.