keep the mass of a species comparatively homogeneous
over any area in which it abounds in individuals.
Starting from a suggestion of the late Mr. Knight,
now so familiar, that close interbreeding diminishes
vigor and fertility; [I-8] and perceiving that bisexuality
is ever aimed at in Nature—being attained
physiologically in numerous cases where it is not
structurally—Mr. Darwin has worked out
the subject in detail, and shown how general is the
concurrence, either habitual or occasional, of two
hermaphrodite individuals in the reproduction of their
kind; and has drawn the philosophical inference that
probably no organic being self-fertilizes indefinitely;
but that a cross with another individual is occasionally—perhaps
at very long intervals—indispensable.
We refer the reader to the section on the intercrossing
of individuals (pp. 96—101), and also to
an article in the Gardeners’ Chronicle a year
and a half ago, for the details of a very interesting
contribution to science, irrespective of theory.
In domestication, this intercrossing may be prevented;
and in this prevention lies the art of producing varieties.
But “the art itself is Nature,” since
the whole art consists in allowing the most universal
of all natural tendencies in organic things (inheritance)
to operate uncontrolled by other and obviously incidental
tendencies. No new power, no artificial force,
is brought into play either by separating the stock
of a desirable variety so as to prevent mixture, or
by selecting for breeders those individuals which
most largely partake of the peculiarities for which
the breed is valued. {I-9]
We see everywhere around us the remarkable results
which Nature may be said to have brought about under
artificial selection and separation. Could she
accomplish similar results when left to herself?
Variations might begin, we know they do begin, in
a wild state. But would any of them be preserved
and carried to an equal degree of deviation?
Is there anything in Nature which in the long-run
may answer to artificial selection? Mr. Darwin
thinks that there is; and Natural Selection is the
key-note of his discourse,
As a preliminary, he has a short chapter to show that
there is variation in Nature, and therefore something
for natural selection to act upon. He readily
shows that such mere variations as may be directly
referred to physical conditions (like the depauperation
of plants in a sterile soil, or their dwarfing as
they approach an Alpine summit, the thicker fur of
an animal from far northward, etc.), and also
those individual differences which we everywhere recognize
but do not pretend to account for, are not separable
by any assignable line from more strongly-marked varieties;
likewise that there is no clear demarkation between
the latter and sub-species, or varieties of the highest
grade (distinguished from species not by any known
inconstancy, but by the supposed lower importance of
their characteristics); nor between these and recognized
species. “These differences blend into
each other in an insensible series, and the series
impresses the mind with an idea of an actual passage.”