modify the received idea of the entire fixity of species.
This field, rich with various but unsystematized stores
of knowledge accumulated by cultivators and breeders,
has been generally neglected by naturalists, because
these races are not in a state of nature; whereas
they deserve particular attention on this very account,
as experiments, or the materials for experiments,
ready to our hand. In domestication we vary some
of the natural conditions of a species, and thus learn
experimentally what changes are within the reach of
varying conditions in Nature. We separate and
protect a favorite race against its foes or its competitors,
and thus learn what it might become if Nature ever
afforded it equal opportunities. Even when, to
subserve human uses, we modify a domesticated race
to the detriment of its native vigor, or to the extent
of practical monstrosity, although we secure forms
which would not be originated and could not be perpetuated
in free Nature, yet we attain wider and juster views
of the possible degree of variation. We perceive
that some species are more variable than others, but
that no species subjected to the experiment persistently
refuses to vary; and that, when it has once begun
to vary, its varieties are not the less but the more
subject to variation. “No case is on record
of a variable being ceasing to be variable under cultivation.”
It is fair to conclude, from the observation of plants
and animals in a wild as well as domesticated state,
that the tendency to vary is general, and even universal.
Mr. Darwin does “not believe that variability
is an inherent and necessary contingency, under all
circumstances, with all organic beings, as some authors
have thought.” No one supposes variation
could occur under all circumstances; but the facts
on the whole imply a universal tendency, ready to be
manifested under favorable circumstances. In
reply to the assumption that man has chosen for domestication
animals and plants having an extraordinary inherent
tendency to vary, and likewise to withstand diverse
climates, it is asked:
“How could a savage possibly know, when he first
tamed an animal, whether it would vary in succeeding
generations and whether it would endure other climates?
Has the little variability of the ass or Guinea-fowl,
or the small power of endurance of warmth by the reindeer,
or of cold by the common camel, prevented their domestication?
I cannot doubt that if other animals and plants, equal
in number to our domesticated productions, and belonging
to equally diverse classes and countries, were taken
from a state of nature, and could be made to breed
for an equal number of generations under domestication,
they would vary on an average as largely as the parent
species of our existing domesticated productions have
varied.”