Variations are an organism’s way of getting over an unexpected discrepancy between its resources as shown by the fly-leaves of its own cheques and the universe’s passbook; the universe is generally right, or would be upheld as right if the matter were to come before the not too incorruptible courts of nature, and in nine cases out of ten the organism has made the error in its own favour, so that it must now pay or die. It can only pay by altering its mode of life, and how long is it likely to be before a new departure in its mode of life comes out in its own person and in those of its family? Granted it will at first come out in their appearance only, but there can be no change in appearance without some slight corresponding organic modification. In practice there is usually compromise in these matters. The universe, if it does not give an organism short shrift and eat it at once, will commonly abate something of its claim; it gets tricked out of an additional moiety by the organism; the organism really does pay something by way of changed habits; this results in variation, in virtue of which the accounts are cooked, cobbled, and passed by a series of those miracles of inconsistency which was call compromises, and after this they cannot be reopened—not till next time.
Surely of the two factors which go to the making up of development, cunning is the one more proper to be insisted on as determining the physical and psychical well or ill being, and hence, ere long, the future form of the organism. We can hardly open a newspaper without seeing some sign of this; take, for example, the following extract from a letter in the Times of the day on which I am writing (February 8, 1886)— “You may pass along a road which divides a settlement of Irish Celts from one of Germans. They all came to the country equally without money, and have had to fight their way in the forest, but the difference in their condition is very remarkable; on the German side there is comfort, thrift, peace, but on the other side the spectacle is very different.” Few will deny that slight organic differences, corresponding to these differences of habit, are already perceptible; no Darwinian will deny that these differences are