of these is the “Portuguese man-of-war,”
the body of which consists of a large pear-shaped
vesicle which floats on the water like a bladder.
From the lower part of this depend into the water
large and small nutritive branches, each ending in
a mouth surrounded by a circle of waving tentacles
armed with batteries of thread-cells, while another
set of hanging protrusions bear the grape-like reproductive
organs. On the upper surface of the bladder is
fixed a purple sail of the most brilliant colour, by
which the floating creature is blown through the water.
When the weather is rough, the bladder empties, and
the creature sinks down into the quiet water below
the waves, to rise again when the storm is over.
This, and its equally wonderful allies, Huxley showed
to be a complicated colony of hydra-like creatures,
each part being composed of two membranes, and therefore
essentially similar to Medusae. Thus, by a great
piece of constructive work, an assemblage of animals
was gathered into a new group and shewn to be organised
upon one simple and uniform plan, and, even in the
most complex and aberrant forms, reducible to the
same type. The group, and Huxley’s conception
of its structure, are now absolutely accepted by anatomists,
and have made one of the corner-stones of our modern
idea of the arrangement of the animal kingdom.
With the exception of sponges, concerning the exact
relations of which there is still dispute, and of a
few sets of parasitic and possibly degenerate creatures,
all animals, the bodies of which are multicellular,
from the simple fresh-water hydra up to man, are divided
into two great groups. The structure of the simpler
of these groups is exactly what Huxley found to be
of importance in the Medusae. The body wall,
from which all the organs protrude, consists merely
of a web of cells arranged in two sheets or membranes,
and the single cavity consists of a central stomach,
surrounded by these membranes, the cavity remaining
simple or giving rise to a number of branching canals.
The members of this great division of the animal kingdom
are the creatures which Huxley selected and placed
together, with the addition of the sea-anemones and
the medusa-like Ctenophora, which, indeed, he mentioned
in his memoir as being related to the others, but
reserved fuller consideration for a future occasion.
This group is now called the Coelenterata, the name
implying that the creatures are simply hollow stomachs,
and it is contrasted in the strongest way with the
group Coelomata, in which are placed all the higher
animals, from the simplest worm up to man; animals
in which, in addition to the two foundation-membranes
of the Coelenterata, there is a third foundation-membrane,
and in which, in addition to the simple stomach cavity
with its offshoots, there is a true body-cavity or
coelome, and usually a set of spaces and channels
containing a blood-fluid. The older method of
naming groups of animals after some obvious superficial
character lingered on for some years in text-books
and treatises, but in this memoir the young ship-surgeon
had replaced it by the modern scientific method of
grouping animals together only because of real identity
of structure.