to pick out the insects as one group and to call everything
else “Vermes” or worms. The insects
included all creatures possessed of an external skeleton
or hard skin divided into jointed segments, and included
forms so different as insects, spiders, crabs, and
lobsters. But Vermes included all the members
of the animal kingdom that were neither vertebrates
nor insects. Cuvier advanced a little. He
got rid of the comprehensive title Vermes—the
label of the rubbish-heap of zooelogists. He
divided animals into four great subkingdoms:
Vertebrates, Mollusca, Articulata, Radiata. These
names, however, only covered very superficial resemblances
among the animals designated by them. The word
Mollusca only meant that the creatures grouped
together had soft bodies, unsupported by internal or
external articulated skeletons; and this character,
or, rather, absence of character, was applied alike
to many totally dissimilar creatures. The term
Articulata included not only Linnaeus’s
insects but a number of soft-skinned, apparently jointed,
worm-like animals such as the leech and earthworm.
Lastly, the name
Radiata meant no more than
that the organs of the creatures so designated were
more or less disposed around a centre, as the sepals
and petals of a flower are grouped around the central
pistil; and it included animals so different as the
starfish and sea-anemones and Medusae. The names
used in the classification were not only loosely applied
but were based on the most superficial observation,
and took no account of the intimate structures of
the tissues and organs of the animals. With slight
modifications, due to individual taste or special knowledge
of small groups, later writers had followed Linnaeus
and Cuvier.
It was with a view of the animal kingdom not much
clearer than this that Huxley began his work on the
Medusae of the tropic seas. He began to study
them no doubt simply because they were among the most
abundant of the animals that could be obtained from
the ship. He made endless dissections and drawings,
and, above all, studied their minute anatomy with
the microscope. They were all placed among Cuvier’s
Radiata, but, as Huxley said in the first line
of his memoir:
“Perhaps no class of animals
has been investigated with so little satisfactory
and comprehensive result, and this not for the want
of patience and ability on the part of the observers,
but rather because they have contented themselves
with stating matters of detail concerning particular
genera and species, instead of giving broad and
general views of the whole class, considered as organised
upon a given type, and inquiring into its relations
with other families.”
He found that fully developed Medusae consisted each
of a disc with tentacles and vesicular bodies at the
margins, a stomach, and canals proceeding from it,
and generative organs. He traced this simple
common structure through the complications and modifications