to the real or mythical conditions of the toad-in-a-hole.
That curious West African mud-fish, the Lepidosiren
(familiar to all readers of evolutionary literature
as one of the most singular existing links between
fish and amphibians), lives among the shallow pools
and broads of the Gambia, which are dried up during
the greater part of the tropical summer. To provide
against this annual contingency, the mud-fish retires
into the soft clay at the bottom of the pools, where
it forms itself a sort of nest, and there hibernates,
or rather aestivates, for months together, in a torpid
condition. The surrounding mud then hardens into
a dry ball; and these balls are dug out of the soil
of the rice-fields by the natives, with the fish inside
them, by which means many specimens of lepidosiren
have been sent alive to Europe, embedded in their natural
covering. Here the strange fish is chiefly prized
as a zoological curiosity for aquariums, because of
its possessing gills and lungs together, to fit it
for its double existence; but the unsophisticated
West Africans grub it up on their own account as a
delicacy, regardless of its claims to scientific consideration
as the earliest known ancestor of all existing terrestrial
animals. Now, the torpid state of the mud-fish
in his hardened ball of clay closely resembles the
real or supposed condition of the toad-in-a-hole;
but with one important exception. The mud-fish
leaves a small canal or pipe open in his cell at either
end to admit the air for breathing, though he breathes
(as I shall proceed to explain) in a very slight degree
during his aestivation; whereas every proper toad-in-a-hole
ought by all accounts to live entirely without either
feeding or breathing in any way. However, this
is a mere detail; and indeed, if toads-in-a-hole do
really exist at all, we must in all probability ultimately
admit that they breathe to some extent, though perhaps
very slightly, during their long immurement.
And this leads us on to consider what in reality hibernation
is. Everybody knows nowadays, I suppose, that
there is a very close analogy between an animal and
a steam-engine. Food is the fuel that makes the
animal engine go; and this food acts almost exactly
as coal does in the artificial machine. But coal
alone will not drive an engine; a free draught of
open air is also required in order to produce combustion.
Just in like manner the food we eat cannot be utilised
to drive our muscles and other organs unless it is
supplied with oxygen from the air to burn it slowly
inside our bodies. This oxygen is taken into the
system, in all higher animals, by means of lungs or
gills. Now, when we are working at all hard,
we require a great deal of oxygen, as most of us have
familiarly discovered (especially if we are somewhat
stout) in the act of climbing hills or running to
catch a train. But when we are doing very little
work indeed, as in our sleeping hours, during which
muscular movement is suspended, and only the general