Some of the ammonites, which belonged to this cuttle-fish group, soon attained a very considerable size; but a shell known as the orthoceras (I wish my subject didn’t compel me to use such very long words, but I am not personally answerable, thank heaven, for the vagaries of modern scientific nomenclature) grew to a bigger size than that of any other fossil mollusk, sometimes measuring as much as six feet in total length. At what date the gigantic cuttles of the present day first began to make their appearance it would be hard to say, for their shell-less bodies are so soft that they could leave hardly anything behind in a fossil state; but the largest known cuttle, measured by Mr. Gabriel, of Newfoundland, was eighty feet in length, including the long arms.
These cuttles are the only invertebrates at all in the running so far as colossal size is concerned, and it will be observed that here the largest modern specimen immeasurably beats the largest fossil form of the same type. I do not say that there were not fossil forms quite as big as the gigantic calamaries of our own time—on the contrary, I believe there were; but if we go by the record alone we must confess that, in the matter of invertebrates at least, the balance of size is all in favour of our own period.
The vertebrates first make their appearance, in the shape of fishes, towards the close of the Silurian period, the second of the great geological epochs. The earliest fish appear to have been small, elongated, eel-like creatures, closely resembling the lampreys in structure; but they rapidly developed in size and variety, and soon became the ruling race in the waters of the ocean, where they maintained their supremacy till the rise of the great secondary saurians. Even then, in spite of the severe competition thus introduced, and still later, in spite of the struggle for life against the huge modern cetaceans (the true monarchs of the recent seas), the sharks continued to hold their own as producers of gigantic forms; and at the present day their largest types probably rank second only to the whales in the whole range of animated nature. There seems no reason to doubt that modern fish, as a whole, quite equal in size the piscine fauna of any previous geological age.
It is somewhat different with the next great vertebrate group, the amphibians, represented in our own world only by the frogs, the toads, the newts, and the axolotls. Here we must certainly with shame confess that the amphibians of old greatly surpassed their degenerate descendants in our modern waters. The Japanese salamander, by far the biggest among our existing newts, never exceeds a yard in length from snout to tail; whereas some of the labyrinthodonts (forgive me once more) of the Carboniferous Epoch must have reached at least seven or eight feet from stem to stern. But the reason of this falling off is not far to seek. When the adventurous newts and frogs of that remote period first dropped their gills and