this latter capacity evidently depends on widely different circumstances—­should all run to a certain extent parallel with the systematic affinity of the forms which are subjected to experiment; for systematic affinity attempts to express all kinds of resemblance between all species.

“First crosses between forms known to be varieties, or sufficiently alike to be considered as varieties, and their mongrel offspring, are very generally, but not quite universally, fertile.  Nor is this nearly general and perfect fertility surprising, when we remember how liable we are to argue in a circle with respect to varieties in a state of Nature; and when we remember that the greater number of varieties have been produced under domestication by the selection of mere external differences, and not of differences in the reproductive system.  In all other respects, excluding fertility, there is a close general resemblance between hybrids and mongrels.”—­Pp. 276-8.

We fully agree with the general tenor of this weighty passage; but forcible as are these arguments, and little as the value of fertility or infertility as a test of species may be, it must not be forgotten that the really important fact, so far as the inquiry into the origin of species goes, is, that there are such things in Nature as groups of animals and of plants, whose members are incapable of fertile union with those of other groups; and that there are such things as hybrids, which are absolutely sterile when crossed with other hybrids.  For if such phaenomena as these were exhibited by only two of those assemblages of living objects, to which the name of species (whether it be used in its physiological or in its morphological sense) is given, it would have to be accounted for by any theory of the origin of species, and every theory which could not account for it would be, so far, imperfect.

Up to this point we have been dealing with matters of fact, and the statements which we have laid before the reader would, to the best of our knowledge, be admitted to contain a fair exposition of what is at present known respecting the essential properties of species, by all who have studied the question.  And whatever may be his theoretical views, no naturalist will probably be disposed to demur to the following summary of that exposition:—­

Living beings, whether animals or plants, are divisible into multitudes of distinctly definable kinds, which are morphological species.  They are also divisible into groups of individuals, which breed freely together, tending to reproduce their like, and are physiological species.  Normally resembling their parents, the offspring of members of these species are still liable to vary, and the variation may be perpetuated by selection, as a race, which race, in many cases, presents all the characteristics of a morphological species.  But it is not as yet proved that a race ever exhibits, when crossed with another race of the same species, those phaenomena of hybridization which are exhibited by many species when crossed with other species.  On the other hand, not only is it not proved that all species give rise to hybrids infertile inter se, but there is much reason to believe that, in crossing, species exhibit every gradation from perfect sterility to perfect fertility.