Palaeozoic period, could we find them, might contain
other orders of vertebrata. But no such reply
can be made to the argument that whereas the marine
vertebrata of the Palaeozoic period consisted entirely
of cartilaginous fishes, the marine vertebrata of
later periods include numerous genera of osseous fishes;
and that, therefore, the later marine vertebrate faunas
are more heterogeneous than the oldest known one.
Nor, again, can any such reply be made to the fact
that there are far more numerous orders and genera
of mammalian remains in the tertiary formations than
in the secondary formations. Did we wish merely
to make out the best case, we might dwell upon the
opinion of Dr. Carpenter, who says that “the
general facts of Palaeontology appear to sanction the
belief, that
the same plan may be traced out
in what may be called
the general life of the globe,
as in
the individual life of every one of the
forms of organised being which now people it.”
Or we might quote, as decisive, the judgment of Professor
Owen, who holds that the earlier examples of each
group of creatures severally departed less widely
from archetypal generality than the later ones—were
severally less unlike the fundamental form common
to the group as a whole; that is to say—constituted
a less heterogeneous group of creatures; and who further
upholds the doctrine of a biological progression.
But in deference to an authority for whom we have
the highest respect, who considers that the evidence
at present obtained does not justify a verdict either
way, we are content to leave the question open.
Whether an advance from the homogeneous to the heterogeneous
is or is not displayed in the biological history of
the globe, it is clearly enough displayed in the progress
of the latest and most heterogeneous creature—Man.
It is alike true that, during the period in which the
Earth has been peopled, the human organism has grown
more heterogeneous among the civilised divisions of
the species; and that the species, as a whole, has
been growing more heterogeneous in virtue of the multiplication
of races and the differentiation of these races from
each other.
In proof of the first of these positions, we may cite
the fact that, in the relative development of the
limbs, the civilised man departs more widely from
the general type of the placental mammalia than do
the lower human races. While often possessing
well-developed body and arms, the Papuan has extremely
small legs: thus reminding us of the quadrumana,
in which there is no great contrast in size between
the hind and fore limbs. But in the European,
the greater length and massiveness of the legs has
become very marked—the fore and hind limbs
are relatively more heterogeneous. Again, the
greater ratio which the cranial bones bear to the
facial bones illustrates the same truth. Among
the vertebrata in general, progress is marked by an
increasing heterogeneity in the vertebral column,
and more especially in the vertebrae constituting the