But, one great service that has been rendered by Mr. Darwin to the doctrine of evolution in general is this: he has shown that there are two chief factors in the process of evolution: one of them is the tendency to vary, the existence of which in all living forms may be proved by observation; the other is the influence of surrounding conditions upon what I may call the parent form and the variations which are thus evolved from it. The cause of the production of variations is a matter not at all properly understood at present. Whether variation depends upon some intricate machinery—if I may use the phrase—of the living organism itself, or whether it arises through the influence of conditions upon that form, is not certain, and the question may, for the present, be left open. But the important point is that granting the existence of the tendency to the production of variations; then, whether the variations which are produced shall survive and supplant the parent, or whether the parent form shall survive and supplant the variations, is a matter which depends entirely on those conditions which give rise to the struggle for existence. If the surrounding conditions are such that the parent form is more competent to deal with them, and flourish in them than the derived forms, then, in the struggle for existence, the parent form will maintain itself and the derived forms will be exterminated. But if, on the contrary, the conditions are such as to be more favourable to a derived than to the parent form, the parent form will be extirpated and the derived form will take its place. In the first case, there will be no progression, no change of structure, through any imaginable series of ages; in the second place there will be modification of change and form.
Thus the existence of these persistent types, as I have termed them, is no real obstacle in the way of the theory of evolution. Take the case of the scorpions to which I have just referred. No doubt, since the Carboniferous epoch, conditions have always obtained, such as existed when the scorpions of that epoch flourished; conditions in which scorpions find themselves better off, more competent to deal with the difficulties in their way, than any variation from the scorpion type which they may have produced; and, for that reason, the scorpion type has persisted, and has not been supplanted by any other form. And there is no reason, in the nature of things, why, as long as this world exists, if there be conditions more favourable to scorpions than to any variation which may arise from them, these forms of life should not persist.
Therefore, the stock objection to the hypothesis of evolution, based on the long duration of certain animal and vegetable types, is no objection at all. The facts of this character—and they are numerous—belong to that class of evidence which I have called indifferent. That is to say, they may afford no direct support to the doctrine of evolution, but they are capable of being interpreted in perfect consistency with it.