He then refers to some of the geological “principles” expounded by Sir Charles Lyell on the “extinction of species,” and follows this up by saying:
To discover how the extinct species have from time to time been replaced by new ones down to the very latest geological period, is the most difficult, and at the same time the most interesting, problem in the natural history of the earth. The present inquiry, which seeks to eliminate from known facts a law which has determined, to a certain degree, what species could and did appear at a given epoch, may, it is hoped, be considered as one step in the right direction towards a complete solution of it.... Admitted facts seem to show ... a general, but not a detailed progression.... It is, however, by no means difficult to show that a real progression in the scale of organisation is perfectly consistent with all the appearances, and even with apparent retrogression should such occur.
Using once more the analogy of a branching tree to illustrate the natural arrangement of species and their successive creation, he clearly shows how “apparent retrogression may be in reality a progress, though an interrupted one”; as “when some monarch of the forest loses a limb, it may be replaced by a feeble and sickly substitute.” As an instance he mentions the Mollusca, which at an early period had reached a high state of development of forms and species, while in each succeeding age modified species and genera replaced the former ones which had become extinct, and “as we approach the present era but few and small representatives of the group remain, while the Gasteropods and Bivalves have acquired an immense preponderance.” In the long series of changes the earth had undergone, the process of peopling it with organic beings had been continually going on, and whenever any of the higher groups had become nearly or quite extinct, the lower forms which better resisted the modified physical conditions served as the antetype on which to found new races. In this manner alone, it was believed, could the representative groups of successive periods, and the risings and fallings in the scale of organisations, be in every case explained.
Again, attending to a recent article by Prof. Forbes, he points out certain inaccuracies and how they may be proved to be so; and continues: