Even in the arbitrary character, which is considered as destructive of all their other claims to ordinary vegetable rank, there is no unison whatever, for Rafflesiaceae have ordinary ovula, while Sarcocoidalis very extraordinary.
The amount of testimony proving their analogy in germination to be with Acrogens, must be very strong before I am convinced that plants with perfect ovula as Rafflesia, etc. germinate from an indeterminate point, the existence of an aperture in the coats, points in the most marked manner to some part representing a radicle. With the exception perhaps of Sarcocoidalis, these plants differ in no respect whatever from other Phaenogamous vegetables; we have instances of the same parasitical growth, and instances of the same apparent want of a radicle or homogeneousness of embryo, and in the structure of the parts of the flower there is tolerably absolute general identity.
It may be worthy of remark, as tending to prove the soundness of Mr. Brown’s views with regard to the affinity of Rafflesia with Aristolochia, that a certain large and fleshy flowered species of the latter genus has the same putrescent smelling flowers.
In Rhizantheae, as proposed by Endlicher, we have an assemblage of discordant characters; we have plants associated, differing in the number of their parts; we have some of comparatively simple roots associated with others of decidedly complex organization; we have Rafflesia in which highly complex female parts exist, associated with Sarcocoidalis, in which these are very simple. But besides the objection of combining discrepancies on the strength of one agreement, the establishment of divisions upon such pretexts is objectionable in another point of view; viz., that of making a transition of structure on one point, instead of in several.
We might as well form into one division all the ternarily formed Dicotyledons, and into another all those Monocotyledonous plants with evident distinction between the calyx and corolla.
But in addition to reasons founded on structure, I have this theoretical one, that it is as requisite that Endogens should establish a similar relation with Acrogens; otherwise a gradation exists between the first and third classes, and none between the second and third, between which, gradations ought to be the more frequent.
As Rafflesia approaches Aristolochia, so does Sarcocodon, Taccaceae.
23rd.—Rawil Pendi. The country continues much the same to within five or six miles of this place, viz. high raviny ground, well covered with Mimosa, Bheir, etc.
Thence to Pendi, the country is open, bare, and much cultivated. From high ground near Pendi a considerable tract is visible, consisting of low ridges running nearly due south, interrupted here and there, and apparently quite bare.
24th.—To Manikyala, distance nineteen miles, over an elevated country, with not much cultivation; broken ground occurs here and there, especially near the river Hoomook, now a small stream, the road winding through Mimosa jungle. Moacurra, Bheir, Euonymus.