The nature of the barren fronds requires distinct analysis.  Are they barren from mere deficiency in supplies, such as may result from many circumstances; or are the antheriform ramenta deficient?  They are barren from defective growth.  I am aware how readily objection may be taken to these views, some will say these young ramenta are nothing but young scales as the older ones evidently are scales; but this amounts to nothing, because we may expect simplicity in the sexual organs of this division, and it will be only a proof of the uniformity of nature in making so great a difference in a function depend on, or be associated with so small a one in form.  My view I think explains their uniformly brown colour—­analogous to Brown’s sphacelation in mutatis mutandis.

Others will say how absurd the idea is, when you cannot show the place to which the impregnating influence is to be applied.  But the consideration of mosses does away with this objection partly, and that of Anthoceros, entirely; because in mosses, the ovule, or pre-existing cell, ready to receive the male influence becomes an empty cell, terminating the seta; and the sporula become developed at its opposite end, the first growth appearing to be quite unconnected with that of the future reproductive organs:  and in Anthoceros there is no fixed punctum ready for the application of the male organs, but these have to form a communication with the lower, or inferior cellular tissue of the frond, before even the growth of seta can commence.

Besides a case in point exists in Viscum, or Loranthus, in which no point is ready prepared for the reception of the male influence; showing how universal the law is, that in no one point or place is there an absolute want of gradation.

As in mosses the influence of the male disregarding the ovule, is thrown into the development of the seta, and then of the theca at the apex of this; there can be no conclusive reason why in ferns the same influence should be thrown into the development of the frond, and then into that of the theca.

While Anthoceros proves that in these orders the male influence may exert its effects upon any point.

As there is no styliform production in Anthoceros, so there is none in ferns.  If the ramenta be anthers, they will not be dubious ones, because as they remain fixed, people cannot say, that possibly they are also reproductive bodies, which by the bye is no objection at all, after instances of anthers bearing ovules instead of pollen!

Why the peculiar distribution of the male influence (on which we determine our genera,) takes place, is another question, and one that cannot be fairly asked?

Why it is confined to the under surface perhaps can, it being a law that in all cases it is the under surface of the leaf, or its modification, from which new growths originate, and as nature has closed indusia, how could the under surface be interior if this rule were not regularly adhered to?