and a few characteristic forms are here figured (fig.
72). Of these, no genus is perhaps more characteristic
than
Euomphalus (fig. 72, b), with its flat
discoidal shell, coiled up into an oblique spiral,
and deeply hollowed out on one side; but examples
of this group are both of older and of more modern
date. Another very extensive genus, especially
in America, is Platyceras (fig. 72, a and f), with
its thin fragile shell—often hardly coiled
up at all—its minute spire, and its widely-expanded,
often sinuated mouth. The British
Acroculioe
should probably be placed here, and the group has
with reason been regarded as allied to the Violet-snails
(
Ianthina) of the open Atlantic. The species
of
Platyostoma (fig. 72, h) also belong to
the same family; and the entire group is continued
throughout the Devonian into the Carboniferous.
Amongst other well-known Upper Silurian Gasteropods
are species of the genera
Holopea (fig. 72,
g),
Holopella (fig. 72. e),
Platyschisma
(fig. 72, d),
Cyclonema, Pleurotomaria, Murchisonia,
Trochonema, &c. The oceanic Univalves (
Heteropods)
are represented mainly by species of
Bellerophon;
and the Winged Snails, or
Pteropods, can still
boast of the gigantic
Thecoe and
Conularioe,
which characterise yet older deposits. The commonest
genus of
Pteropoda, however, is
Tentaculites
(fig. 73), which clearly belongs here, though it has
commonly been regarded as the tube of an Annelide.
The shell in this group is a conical tube, usually
adorned with prominent transverse rings, and often
with finer transverse or longitudinal striae as well;
and many beds of the Upper Silurian exhibit myriads
of such tubes scattered promiscuously over their surfaces.
The last and highest group of the Mollusca—that
of the Cephalopoda—is still represented
only by Tetrabranchiate forms; but the abundance
and variety of these is almost beyond belief.
Many hundreds of different species are known, chiefly
belonging to the straight Orthoceratites, but
the slightly-curved Cyrtoceras is only little
less common. There are also numerous forms of
the genera Phragmoceras, Ascoceras, Gyroteras, Lituites,
and Nautilus. Here, also, are the first-known
species of the genus Goniatites—a
group which attains considerable importance in later
deposits, and which is to be regarded as the precursor
of the Ammonites of the Secondary period.
[Illustration: Fig. 74.—Head-shield
of Pteraspis Banksii, Ludlow rocks. (After
Murchison.)]
[Illustration: Fig. 75.—A, Spine of
Onchus tenuistriatus; B, Shagreen-scales of
Thelodus. Both from the “bone-bed”
of the Upper Ludlow rocks. (After Murchison.)]