structureless animal substance of an albuminous nature
("sarcode"), of a gelatinous consistence, transparent,
and exhibiting numerous minute granules or rounded
particles. The body-substance cannot be said
in itself to possess any definite form, except in
so far as it may be bounded by a shell; but it has
the power, wherever it may be exposed, of emitting
long thread-like filaments ("pseudopodia"), which
interlace with one another to form a network (fig.
25, b). These filaments can be thrown out at
will, and to considerable distances, and can be again
retracted into the soft mass of the general body-substance,
and they are the agents by which the animal obtains
its food. The soft bodies of the
Foraminifera
are protected by a shell, which is usually calcareous,
but may be composed of sand-grains cemented together;
and it may consist of a single chamber (fig. 26, a),
or of many chambers arranged in different ways (fig.
26,
b-f). Sometimes the shell has but one
large opening into it—the mouth; and then
it is from this aperture that the animal protrudes
the delicate net of filaments with which it seeks its
food. In other cases the entire shell is perforated
with minute pores (fig. 26, e), through which the
soft body-substance gains the exterior, covering the
whole shell with a gelatinous film of animal matter,
from which filaments can be emitted at any point.
When the shell consists of many chambers, all of these
are placed in direct communication with one another,
and the actual substance of the shell is often traversed
by minute canals filled with living matter (e.g.,
in
Calcarina and
Nummulina). The
shell, therefore, may be regarded, in such cases, as
a more or less completely porous calcareous structure,
filled to its minutest internal recesses with the
substance of the living animal, and covered externally
with a layer of the same substance, giving off a network
of interlacing filaments.
[Illustration: Fig. 25.—The animal
of Nonionina, one of the Foraminifera,
after the shell has been removed by a weak acid; b,
Gromia, a single-chambered Foraminifer (after
Schultze), showing the shell surrounded by a network
of filaments derived from the body substance.]
[Illustration: Fig 26.—Shells of living
Foraminifera. a, Orbulina universa,
in its perfect condition, showing the tubular spines
which radiate from the surface of the shell; b, Globigerina
bulloides, in its ordinary condition, the thin
hollow spines which are attached to the shell when
perfect having been broken off; c, Textularia variabilis;
d, Peneroplis planatus; e, Rotalia concamerata; f,
Cristellaria subarcuatula. [Fig. a is after
Wyville Thomson; the others are after Williamson.
All the figures are greatly enlarged.]]