(fig. 23, a a). This proper wall forms the actual
lining of the chambers, as well as the outer surface
of the whole mass; and it is perforated with numerous
fine vertical tubes (fig. 24, a a), opening into the
chambers and on to the surface by corresponding fine
pores. From the resemblance of this tubulated
layer to similar structures in the shell of the Nummulite,
it is often spoken of as the “Nummuline layer.”
The chambers are sometimes piled up one above the
other in an irregular manner; but they are more commonly
arranged in regular tiers, the separate chambers being
marked off from one another by projections of the
wall in the form of partitions, which are so far imperfect
as to allow of a free communication between contiguous
chambers. In the original condition of the organism,
all these chambers, of course, must have been filled
with living-matter; but they are found in the present
state of the fossil to be generally filled with some
silicate, such as serpentine, which not only fills
the actual chambers, but has also penetrated the minute
tubes of the proper wall and the branching canals
of the intermediate skeleton. In some cases the
chambers are simply filled with crystalline carbonate
of lime. When the originally porous fossil has
been permeated by a silicate, it is possible to dissolve
away the whole of the calcareous skeleton by means
of acids, leaving an accurate and beautiful cast of
the chambers and the tubes connected with them in
the insoluble silicate.
[Illustration: Fig. 24.—Portion of
one of the calcareous layers of Eozooen, magnified
100 diameters. a a, The proper wall ("Nummuline layer”)
of one of the chambers, showing the fine vertical
tubuli with which it is penetrated, and which are slightly
bent along the line a’ a’. c c, The intermediate
skeleton, with numerous branched canals. The
oblique lines are the cleavage planes of the carbonate
of lime, extending across both the intermediate skeleton
and the proper wall. (After Carpenter.)]
The above are the actual appearances presented by
Eozooen when examined microscopically, and
it remains to see how far they enable us to decide
upon its true position in the animal kingdom.
Those who wish to study this interesting subject in
detail must consult the admirable memoirs by Dr W.
B. Carpenter and Principal Dawson: it will be
enough here to indicate the results which have been
arrived at. The only animals at the present day
which possess a continuous calcareous skeleton, perforated
by pores and penetrated by canals, are certain organisms
belonging to the group of the Foraminifera.
We have had occasion before to speak of these animals,
and as they are not conspicuous or commonly-known
forms of life, it may be well to say a few words as
to the structure of the living representatives of the
group. The Foraminifera are all inhabitants
of the sea, and are mostly of small or even microscopic
dimensions. Their bodies are composed of an apparently