in a horny beak. The breast-bone is entirely
destitute of a central ridge or keel, and the wings
are minute and quite rudimentary; so that
Hesperornis,
unlike
Ichthyornis, must have been wholly deprived
of the power of flight, in this respect approaching
the existing Penguins. The tail consists of about
twelve vertebrae, of which the last three or four
are amalgamated to form a flat terminal mass, there
being at the same time clear indications that the
tail was capable of up and down movement in a vertical
plane, this probably fitting it to serve as a swimming-paddle
or rudder. The legs were powerfully constructed,
and the feet were adapted to assist the bird in rapid
motion through the water. The known remains of
Hesperornis regalis prove it to have been a
swimming and diving bird, of larger dimensions than
any of the aquatic members of the class of Birds with
which we are acquainted at the present day. It
appears to have stood between five and six feet high,
and its inability to fly is fully compensated for by
the numerous adaptations of its structure to a watery
life. Its teeth prove it to have been carnivorous
in its habits, and it probably lived upon fishes.
It is a curious fact that two Birds agreeing with
one another in the wholly abnormal character of possessing
teeth, and in other respects so entirely different,
should, like
Ichthyornis and
Hesperornis,
have lived not only in the same geological period,
but also in the same geographical area; and it is
equally curious that the area inhabited by these toothed
Birds should at the same time have been tenanted by
winged and bird-like Reptiles belonging to the toothed
genus
Pterodactylus and the toothless genus
Pteranodon.
[Illustration: Fig. 212.—Toothed Birds
(Odontornithes) of the Cretaceous Rocks of
America. a. Left lower jaw of Ichthyornis
dispar, slightly enlarged; b, Left lower jaw of
Hesperornis regalis, reduced to nearly one-fourth
of the natural size; c. Cervical vertebra of
Ichthyornis dispar, front view, twice the natural
size; c’, Side view of the same; d, Tooth of
Hesperornis regalis, enlarged to twice the natural
size. (After Marsh.)]
No remains of Mammals, finally, have as yet
been detected in any sedimentary accumulations of
Cretaceous age.
LITERATURE.
The following list comprises some of the more important
works and memoirs which may be consulted with reference
to the Cretaceous strata and their fossil contents:—
(1) ‘Memoirs of the Geological Survey
of Great Britain.’
(2) ‘Geology of England and Wales.’
Conybeare and Phillips.
(3) ‘Geology of Yorkshire,’ vol.
ii. Phillips.
(4) ‘Geology of Oxford and the Thames
Valley.’ Phillips.
(5) ‘Geological Excursions through the
Isle of Wight.’ Mantell.
(6) ‘Geology of Sussex.’ Mantell.
(7) ‘Report on Londonderry,’ &c.