belonging to it, have been discovered in the Cretaceous
deposits of North America, and have been described
by Professors Cope and Marsh. All the known forms
of this group appear to have been of large size—one
of them,
Mosasaurus princeps, attaining the
length of seventy-five or eighty feet, and thus rivalling
the largest of existing Whales in its dimensions.
The teeth in the “Mosasauroids” are long,
pointed, and slightly curved; and instead of being
sunk in distinct sockets, they are firmly amalgamated
with the jaws, as in modern Lizards. The palate
also carried teeth, and the lower jaw was so constructed
as to allow of the mouth being opened to an immense
width, somewhat as in the living Serpents. The
body was long and snake-like, with a very long tail,
which is laterally compressed, and must have served
as a powerful swimming-apparatus. In addition
to this, both pairs of limbs have the bones connecting
them with the trunk greatly shortened; whilst the
digits were enclosed in the integuments, and constituted
paddles, closely resembling in structure the “flippers”
of Whales and Dolphins. The neck is sometimes
moderately long, but oftener very short, as the great
size and weight of the head would have led one to
anticipate. Bony plates seem in some species
to have formed an at any rate partial covering to
the skin; but it is not certain that these integumentary
appendages were present in all. Upon the whole,
there can be no doubt but that the Mosasauroid Reptiles—the
true “Sea-serpents” of the Cretaceous
period—were essentially aquatic in their
habits, frequenting the sea, and only occasionally
coming to the land.
[Illustration: Fig. 210.—Skull of
Mosasaurus Camperi, greatly reduced. Maestricht
Chalk.]
The “Mosasauroids” have generally been
regarded as a greatly modified group of the Lizards
(Lacertilia). Whether this reference be
correct or not—and recent investigations
render it dubious—the Cretaceous rocks
have yielded the remains of small Lizards not widely
removed from existing forms. The recent order
of the Chelonians is also represented in the
Cretaceous rocks, by forms closely resembling living
types. Thus the fresh-water deposits of the Wealden
have yielded examples of the “Terrapins”
or “Mud-Turtles” (Emys); and the
marine Cretaceous strata have been found to contain
the remains of various species of Turtles, one of which
is here figured (fig. 211). No true Serpents (Ophidia)
have as yet been detected in the Cretaceous rocks;
and this order does not appear to have come into existence
till the Tertiary period. Lastly, true Crocodiles
are known to have existed in considerable numbers
in the Cretaceous period. The oldest of these
occur in the fresh-water deposit of the Wealden; and
they differ from the existing forms of the group in
the fact that the bodies of the vertebrae, like those
of the Jurassic Crocodiles, are bi-concave, or hollowed
out at both ends. In the Greensand of North America,
however, occur the remains of Crocodiles which agree
with all the living species in having the bodies of
the vertebrae in the region of the back hollowed out
in front and convex behind.