the jaw, instead of being sunk in distinct sockets;
and they have the form of somewhat flattened prisms,
longitudinally ridged on the outer surface, with an
obtusely triangular crown, and having the enamel crenated
on one or both sides. They present the extraordinary
feature that the crowns became worn down flat by mastication,
showing that the Iguanodon employed its teeth
in actually chewing and triturating the vegetable matter
on which it fed. There can therefore be no doubt
but that the Iguanodon, in spite of its immense
bulk, was an herbivorous Reptile, and lived principally
on the foliage of the Cretaceous forests amongst which
it dwelt. Its size has been variously estimated
at from thirty to fifty feet, the thigh-bone in large
examples measuring nearly five feet in length, with
a circumference of twenty-two inches in its smallest
part. With the strong and massive hind-limbs are
associated comparatively weak and small fore-limbs;
and there seems little reason to doubt that the Iguanodon
must have walked temporarily or permanently upon its
hind-limbs, after the manner of a Bird. This
conjecture is further supported by the occurrence in
the strata which contain the bones of the Iguanodon
of gigantic three-toed foot-prints, disposed singly
in a double track. These prints have undoubtedly
been produced by some animal walking on two legs;
and they can hardly, with any probability, be ascribed
to any other than this enormous Reptile. Closely
allied to the Iguanodon is the Hadrosaurus
of the American Cretaceous, the length of which is
estimated at twenty-eight feet. Iguanodon does
not appear to have possessed any integumentary skeleton;
but the great Hyloeosaurus of the Wealden seems
to have been furnished with a longitudinal crest of
large spines running down the back, similar to that
which is found in the comparatively small Iguanas
of the present day. The Megalosaurus of
the Oolites continued to exist in the Cretaceous period;
and, as we have previously seen, it was carnivorous
in its habits. The American Loelaps was
also carnivorous, and, like the Megalosaur, which it
very closely resembles, appears to have walked upon
its hind-legs, the fore-limbs being disproportionately
small.
[Illustration: Fig. 209.—Teeth of Iguanodon Mantellii. Wealden, Britain.]
Another remarkable group of Reptiles, exclusively confined to the Cretaceous series, is that of the Mosasauroids, so called from the type-genus Mosasaurus. The first species of Mosasaurus known to science was the M. Camperi (fig. 210), the skull of which—six feet in length—was discovered in 1780 in the Maestricht Chalk at Maestricht. As this town stands on the river Meuse, the name of Mosasaurus ("Lizard of the Meuse”) was applied to this immense Reptile. Of late years the remains of a large number of Reptiles more or less closely related to Mosasaurus, or absolutely