their apparent total absence in Palaeozoic strata,
is a strong presumption in favour of the view that
the order of the Cuttle-fishes did not come into existence
till the commencement of the Mesozoic period.
The great majority of the skeletons of this kind which
are found in the Jurassic rocks belong to the great
extinct family of the “Belemnites” (
Belemnitidoa),
which, so far as known, is entirely confined to rocks
of Secondary age. From its pointed, generally
cylindro-conical form, the skeleton of the Belemnite
is popularly known as a “thunderbolt”.
(fig. 173, C). In its perfect condition—in
which it is, however, rarely obtainable—the
skeleton consists of a chambered conical shell (the
“phragmacone"), the partitions between the chambers
of which are pierced by a marginal tube or “siphuncle.”
This conical shell—curiously similar in
its structure to the
external shell of the
Nautilus—is extended forwards into a horny
“pen,” and is sunk in a corresponding conical
pit (fig. 173, B), excavated in the substance of a
nearly cylindrical fibrous body or “guard,”
which projects backwards for a longer or shorter distance,
and is the part most usually found in a fossil condition.
Many different kinds of
Belemnites are known,
and their guards literally swarm in many parts of
the Jurassic series, whilst some specimens attain
very considerable dimensions. Not only is the
internal skeleton known, but specimens of
Belemnites
and the nearly allied
Belemnoteuthis have been
found in some of the fine-grained sediments of the
Jurassic formation, from which much has been learnt
even as to the anatomy of the soft parts of the animal.
Thus we know that the Belemnites were in many respects
comparable with the existing Calamaries or Squids,
the body being furnished with lateral fins, and the
head carrying a circle of ten “arms,”
two of which were longer than the others (fig. 173,
A). The suckers on the arms were provided, further,
with horny hooks; there was a large ink-sac; and the
mouth was armed with horny mandibles resembling in
shape the beak of a parrot.
[Illustration: Fig. 173.—A, Restoration
of the animal of the Belemnite; B, Diagram showing
the complete skeleton of a Belemnite, consisting of
the chambered phragmacone (a), the guard (b), and
the horny pen (c); C, Specimen of Belemnites canaliculatus,
from the Inferior Oolite. (After Phillips.)]
[Illustration: Fig. 174.—Tetragonolepis
(restored), and scales of the same. Lias.]
Coming next to the Vertebrates, we find that
the Jurassic Fishes are still represented by
Ganoids and Placoids. The Ganoids,
however, unlike the old forms, now for the most part
possess nearly or quite symmetrical ("homocercal”)
tails. A characteristic genus is Tetragonolepis
(fig. 174), with its deep compressed body, its rhomboidal,
closely-fitting scales, and its single long dorsal
fin. Amongst the Placoids the teeth of