Amongst the Wing-shells (Pteropoda), it is sufficient to mark the final appearance in the Lias of the ancient genus Conularia.
[Illustration: Fig. 170.—Ammonites Humphresianus. Inferior Oolite.]
[Illustration: Fig. 171.—Ammonites bifrons. Lias.]
Lastly, the order of the Cephalopoda, in both its Tetrabranchiate and Dibranchiate sections, undergoes a vast development in the Jurassic period. The old and comparatively simple genus Nautilus is still well represented, one species being very similar to the living Pearly Nautilus (N. Pompilius); but the Orthocerata and Goniatites of the Trias have finally disappeared; and the great majority of the Tetrabranchiate forms are referable to the comprehensive genus Ammonites, with its many sub-genera and its hundreds of recorded species. The shell in Ammonites is in the form of a flat spiral, all the coils of which are in contact (figs. 170 and 171). The innermost whorls of the shell are more or less concealed; and the body-chamber is elongated and narrow, rather than expanded towards the mouth. The tube or siphuncle which runs through the air-chambers is placed on the dorsal or convex side of the shell; but the principal character which distinguishes Ammonites from Goniatites and Ceratites is the wonderfully complex manner in which the septa, or partitions between the air-chambers, are folded and undulated. To such an extent does this take place, that the edges of the septa, when exposed by the removal of the shell-substance, present in an exaggerated manner the appearance exhibited by an elaborately-dressed shirt-frill when viewed edgewise. The species of Ammonites range from the Carboniferous to the Chalk; but they have not been found in deposits older than the Secondary, in any region except India; and they are therefore to be regarded as essentially Mesozoic fossils.