These were the lords of the forests all over the world in the Carboniferous period. Wherever the Carboniferous deposits have been traced, in the United States, in Canada, in England, France, Belgium, Germany, in New Holland, at the Cape of Good Hope, and in South America, the general aspect of the vegetation has been found to be the same, though characterized in the different localities by specific differences of the same nature as those by which the various floras are distinguished now in different parts of the same zone. For instance, the Temperate Zone throughout the world is characterized by certain families of trees: by Oaks, Maples, Beeches, Birches, Pines, etc.; but the Oaks, Maples, Beeches, Birches, and the like, of the American flora in that latitude differ in species from the corresponding European flora. So in the Carboniferous period, when more uniform climatic conditions prevailed throughout the world, the character of the vegetation showed a general unity of structure everywhere; but it was nevertheless broken up into distinct botanical provinces by specific differences of the same kind as those which now give such diversity of appearance to the vegetation of the Temperate Zone in Europe as compared with that of America, or to the forests of South America as compared with those of Africa.
There can be no doubt as to the true nature of the Carboniferous forests; for the structural character of the trees is as strongly marked in their fossil remains as in any living plants of the same character. We distinguish the Ferns not only by the peculiar form of their leaves, often perfectly preserved, but also by the fructification on the lower surface of the leaves, and by the distinct marks made on the stem at their point of juncture with it. The leaf of the Fern, when falling, leaves a scar on the stem varying in shape and size according to the kind of Fern, so that the botanist readily distinguishes any particular species of Fern by this means,—a birth-mark, as it were, by which he detects the parentage of the individual. Another indication, equally significant, is found in the tubular structure of the wood in Ferns. On a vertical section of any well-preserved Fern-trunk from the old forests the little tubes may be seen very distinctly running up its length; or, if it be cut through transversely, they may be traced by the little pores like dots on the surface. Trees of this description are found in the Carboniferous marshes, standing erect and perfectly preserved, with trunks a foot and a half in diameter, rising to a height of many feet. Plants so strongly bituminous as the Ferns, when they equalled in size many of our present forest-trees, naturally made coal deposits of the most combustible quality. It is true that we find the anthracite coal of the same period with comparatively little bituminous matter; but this is where the bitumen has been destroyed by the action of the internal heat of the earth.