I see Professor S. H. Vines, in the article on Weismann’s theory referred to in my preceding article, says Mr. Darwin “held that it was not the sudden variations due to altered external conditions which become permanent, but those slowly produced by what he termed ‘the accumulative action of changed conditions of life.’” Nothing can be more soundly Lamarckian, and nothing should more conclusively show that, whatever else Mr. Darwin was, he was not a Charles-Darwinian; but what evidence other than inferential can from the nature of the case be adduced in support of this, as I believe, perfectly correct judgment? None know better than they who clamour for direct evidence that their master was right in taking the position assigned to him by Professor Vines, that they cannot reasonably look for it. With us, as with themselves, modification proceeds very gradually, and it violates our principles as much as their own to expect visible permanent progress, in any single generation, or indeed in any number of generations of wild species which we have yet had time to observe. Occasionally we can find such cases, as in that of Branchipus stagnalis, quoted by Mr. Wallace, or in that of the New Zealand Kea whose skin, I was assured by the late Sir Julius von Haast, has already been modified as a consequence of its change of food. Here we can show that in even a few generations structure is modified under changed conditions of existence, but as we believe these cases to occur comparatively rarely, so it is still more rarely that they occur when and where we can watch them. Nature is eminently conservative, and fixity of type, even under considerable change of conditions, is surely more important for the well-being of any species than an over-ready power of adaptation to, it may be, passing changes. There could be no steady progress if each generation were not mainly bound by the traditions of those that have gone before it. It is evolution and not incessant revolution that both parties are upholding; and this being so, rapid visible modification must be the exception, not the rule. I have quoted direct evidence adduced by competent observers, which is, I believe, sufficient to establish the fact that offspring can be and is sometimes modified by the acquired habits of a progenitor. I will now proceed to the still more, as it appears to me, cogent proof afforded by general considerations.