Besides, who has seen the uncles and aunts going away with the uniformity that is necessary for Mr. Darwin’s contention? We know that birds and insects do often get blown out to sea and perish, but in order to establish Mr. Darwin’s position we want the evidence of those who watched the reduction of the wings during the many generations in the course of which it was being effected, and who can testify that all, or the overwhelming majority, of the beetles born with fairly well-developed wings got blown out to sea, while those alone survived whose wings were congenitally degenerate. Who saw them go, or can point to analogous cases so conclusive as to compel assent from any equitable thinker?
Darwinians of the stamp of Mr. Thiselton Dyer, Professor Ray Lankester, or Mr. Romanes, insist on their pound of flesh in the matter of irrefragable demonstration. They complain of us for not bringing forward someone who has been able to detect the movement of the hour-hand of a watch during a second of time, and when we fail to do so, declare triumphantly that we have no evidence that there is any connection between the beating of a second and the movement of the hour-hand. When we say that rain comes from the condensation of moisture in the atmosphere, they demand of us a rain-drop from moisture not yet condensed. If they stickle for proof and cavil on the ninth part of a hair, as they do when we bring forward what we deem excellent instances of the transmission of an acquired characteristic, why may not we, too, demand at any rate some evidence that the unmodified beetles actually did always, or nearly always, get blown out to sea, during the reduction above referred to, and that it is to this fact, and not to the masterly inactivity of their fathers and mothers, that the Madeira beetles owe their winglessness? If we begin stickling for proof in this way, our opponents would not be long in letting us know that absolute proof is unattainable on any subject, that reasonable presumption is our highest certainty, and that crying out for too much evidence is as bad as accepting too little. Truth is like a photographic sensitized plate, which is equally ruined by over and by under exposure, and the just exposure for which can never be absolutely determined.
Surely if disuse can be credited with the vast powers involved in Mr. Darwin’s statement that it has probably “been the main agent in rendering organs rudimentary,” no limits are assignable to the accumulated effects of habit, provided the effects of habit, or use and disuse, are supposed, as Mr. Darwin supposed them, to be inheritable at all. Darwinians have at length woke up to the dilemma in which they are placed by the manner in which Mr. Darwin tried to sit on the two stools of use and disuse, and natural selection of accidental variations, at the same time. The knell of Charles-Darwinism is rung in Mr. Wallace’s present book, and in the general perception on the part of biologists that