The trees of the coal forests present parallel conditions. When the fallen trunks which have entered into the composition of the bed of coal are identifiable, they are mere double shells of bark, flattened together in consequence of the destruction of the woody core; and Sir Charles Lyell and Principal Dawson discovered, in the hollow stools of coal trees of Nova Scotia, the remains of snails, millipedes, and salamander-like creatures, embedded in a deposit of a different character from that which surrounded the exterior of the trees. Thus, in endeavouring to comprehend the formation of a seam of coal, we must try to picture to ourselves a thick forest, formed for the most part of trees like gigantic club-mosses, mares-tails, and tree ferns, with here and there some that had more resemblance to our existing yews and fir-trees. We must suppose that, as the seasons rolled by, the plants grew and developed their spores and seeds; that they shed these in enormous quantities, which accumulated on the ground beneath; and that, every now and then, they added a dead frond or leaf; or, at longer intervals, a rotten branch, or a dead trunk, to the mass.
A certain proportion of the spores and seeds no doubt fulfilled their obvious function, and, carried by the wind to unoccupied regions, extended the limits of the forest; many might be washed away by rain into streams, and be lost; but a large portion must have remained, to accumulate like beech-mast, or acorns, beneath the trees of a modern forest.
But, in this case, it may be asked, why does not our English coal consist of stems and leaves to a much greater extent than it does? What is the reason of the predominance of the spores and spore-cases in it?
A ready answer to this question is afforded by the study of a living full-grown club-moss. Shake it upon a piece of paper, and it emits a cloud of fine dust, which falls over the paper, and is the well-known Lycopodium powder. Now this powder used to be, and I believe still is, employed for two objects, which seem at first sight to have no particular connection with one another. It is, or was, employed in making lightning, and in making pills. The coats of the spores contain so much resinous matter, that a pinch of Lycopodium powder, thrown through the flame of a candle, burns with an instantaneous flash, which has long done duty for lightning on the stage. And the same character makes it a capital coating for pills; for the resinous powder prevents the drug from being wetted by the saliva, and thus bars the nauseous flavour from the sensitive papillae of the tongue.
But this resinous matter, which lies in the walls of the spores and sporangia, is a substance not easily altered by air and water, and hence tends to preserve these bodies, just as the bituminized cerecloth preserves an Egyptian mummy; while, on the other hand, the merely woody stem and leaves tend to rot, as fast as the wood of the mummy’s coffin has rotted. Thus the mixed heap of spores, leaves, and stems in the coal-forest would be persistently searched by the long-continued action of air and rain; the leaves and stems would gradually be reduced to little but their carbon, or, in other words, to the condition of mineral charcoal in which we find them; while the spores and sporangia remained as a comparatively unaltered and compact residuum.