their own homes, and are most widely diffused, having produced the greatest number of new varieties.  It is also natural that the dominant, varying, and far-spreading species, which already have invaded to a certain extent the territories of other species, should be those which would have the best chance of spreading still further, and of giving rise in new countries to new varieties and species.  The process of diffusion may often be very slow, being dependent on climatal and geographical changes, or on strange accidents, but in the long run the dominant forms will generally succeed in spreading.  The diffusion would, it is probable, be slower with the terrestrial inhabitants of distinct continents than with the marine inhabitants of the continuous sea.  We might therefore expect to find, as we apparently do find, a less strict degree of parallel succession in the productions of the land than of the sea.

Dominant species spreading from any region might encounter still more dominant species, and then their triumphant course, or even their existence, would cease.  We know not at all precisely what are all the conditions most favourable for the multiplication of new and dominant species; but we can, I think, clearly see that a number of individuals, from giving a better chance of the appearance of favourable variations, and that severe competition with many already existing forms, would be highly favourable, as would be the power of spreading into new territories.  A certain amount of isolation, recurring at long intervals of time, would probably be also favourable, as before explained.  One quarter of the world may have been most favourable for the production of new and dominant species on the land, and another for those in the waters of the sea.  If two great regions had been for a long period favourably circumstanced in an equal degree, whenever their inhabitants met, the battle would be prolonged and severe; and some from one birthplace and some from the other might be victorious.  But in the course of time, the forms dominant in the highest degree, wherever produced, would tend everywhere to prevail.  As they prevailed, they would cause the extinction of other and inferior forms; and as these inferior forms would be allied in groups by inheritance, whole groups would tend slowly to disappear; though here and there a single member might long be enabled to survive.

Thus, as it seems to me, the parallel, and, taken in a large sense, simultaneous, succession of the same forms of life throughout the world, accords well with the principle of new species having been formed by dominant species spreading widely and varying; the new species thus produced being themselves dominant owing to inheritance, and to having already had some advantage over their parents or over other species; these again spreading, varying, and producing new species.  The forms which are beaten and which yield their places to the new and victorious forms, will generally be allied in groups, from inheriting some inferiority in common; and therefore as new and improved groups spread throughout the world, old groups will disappear from the world; and the succession of forms in both ways will everywhere tend to correspond.