parts tend to vary in the same way, and homologous
parts tend to cohere. Modifications in hard parts
and in external parts sometimes affect softer and internal
parts. When one part is largely developed, perhaps
it tends to draw nourishment from the adjoining parts;
and every part of the structure which can be saved
without detriment to the individual, will be saved.
Changes of structure at an early age will generally
affect parts subsequently developed; and there are
very many other correlations of growth, the nature
of which we are utterly unable to understand.
Multiple parts are variable in number and in structure,
perhaps arising from such parts not having been closely
specialised to any particular function, so that their
modifications have not been closely checked by natural
selection. It is probably from this same cause
that organic beings low in the scale of nature are
more variable than those which have their whole organisation
more specialised, and are higher in the scale.
Rudimentary organs, from being useless, will be disregarded
by natural selection, and hence probably are variable.
Specific characters—that is, the characters
which have come to differ since the several species
of the same genus branched off from a common parent—are
more variable than generic characters, or those which
have long been inherited, and have not differed within
this same period. In these remarks we have referred
to special parts or organs being still variable, because
they have recently varied and thus come to differ;
but we have also seen in the second Chapter that the
same principle applies to the whole individual; for
in a district where many species of any genus are
found—that is, where there has been much
former variation and differentiation, or where the
manufactory of new specific forms has been actively
at work—there, on an average, we now find
most varieties or incipient species. Secondary
sexual characters are highly variable, and such characters
differ much in the species of the same group.
Variability in the same parts of the organisation has
generally been taken advantage of in giving secondary
sexual differences to the sexes of the same species,
and specific differences to the several species of
the same genus. Any part or organ developed to
an extraordinary size or in an extraordinary manner,
in comparison with the same part or organ in the allied
species, must have gone through an extraordinary amount
of modification since the genus arose; and thus we
can understand why it should often still be variable
in a much higher degree than other parts; for variation
is a long-continued and slow process, and natural
selection will in such cases not as yet have had time
to overcome the tendency to further variability and
to reversion to a less modified state. But when
a species with any extraordinarily-developed organ
has become the parent of many modified descendants—which
on my view must be a very slow process, requiring a
long lapse of time—in this case, natural