showed wide variation, no two being alike. The
pumas were for the most part bright red, but some
were reddish gray, there being much the same dichromatism
that I found among their Colorado kinsfolk. The
jaguarundis were dark brownish gray. All these
animals, the spotted jaguars and ocelots, the monochrome
black jaguars, red pumas, and dark-gray jaguarundis,
were killed in the same locality, with the same environment.
A glance at the skins and a moment’s serious
thought would have been enough to show any sincere
thinker that in these cats the coloration pattern,
whether concealing or revealing, is of no consequence
one way or the other as a survival factor. The
spotted patterns conferred no benefit as compared with
the nearly or quite monochrome blacks, reds, and dark
grays. The bodily condition of the various beasts
was equally good, showing that their success in life,
that is, their ability to catch their prey, was unaffected
by their several color schemes. Except white,
there is no color so conspicuously advertising as
black; yet the black jaguar had been a fine, well-fed,
powerful beast. The spotted patterns in the forests,
and perhaps even in the marshes which the jaguars so
frequently traversed, are probably a shade less conspicuous
than the monochrome red and gray, but the puma and
jaguarundi are just as hard to see, and evidently
find it just as easy to catch prey, as the jaguar and
ocelot. The little fawn which we saw was spotted;
the grown deer had lost the spots; if the spots do
really help to conceal the wearer, it is evident that
the deer has found the original concealing coloration
of so little value that it has actually been lost in
the course of the development of the species.
When these big cats and the deer are considered, together
with the dogs, tapirs, peccaries, capybaras, and big
ant-eaters which live in the same environment, and
when we also consider the difference between the young
and the adult deer and tapirs (both of which when
adult have substituted a complete or partial monochrome
for the ancestral spots and streaks), it is evident
that in the present life and in the ancestral development
of the big mammals of South America coloration is
not and has not been a survival factor; any pattern
and any color may accompany the persistence and development
of the qualities and attributes which are survival
factors. Indeed, it seems hard to believe that
in their ordinary environments such color schemes
as the bright red of the marsh-deer, the black of
the black jaguar, and the black with white stripes
of the great tamandua, are not positive detriments
to the wearers. Yet such is evidently not the
case. Evidently the other factors in species-survival
are of such overwhelming importance that the coloration
becomes negligible from this standpoint, whether it
be concealing or revealing. The cats mould themselves
to the ground as they crouch or crawl. They take
advantage of the tiniest scrap of cover. They
move with extraordinary stealth and patience.