A certain proportion of the spores and seeds no doubt fulfilled their obvious function, and, carried by the wind to unoccupied regions, extended the limits of the forest; many might be washed away by rain into streams, and be lost; but a large portion must have remained, to accumulate like beech-mast, or acorns, beneath the trees of a modern forest.
But, in this case it may be asked, why does not our English coal consist of stems and leaves to a much greater extent than it does? What is the reason of the predominance of the spores and spore-cases in it?
A ready answer to this question is afforded by the study of a living full-grown club-moss. Shake it upon a piece of paper, and it emits a cloud of fine dust, which falls over the paper, and is the well-known Lycopodium powder. Now this powder used to be, and I believe still is, employed for two objects which seem, at first sight, to have no particular connection with one another. It is, or was, employed in making lightning, and in making pills. The coats of the spores contain so much resinous matter, that a pinch of Lycopodium powder, thrown through the flame of a candle, burns with an instantaneous flash, which has long done duty for lightning on the stage. And the same character makes it a capital coating for pills; for the resinous powder prevents the drug from being wetted by the saliva, and thus bars the nauseous flavour from the sensitive papilla; of the tongue.
But this resinous matter, which lies in the walls of the spores and sporangia, is a substance not easily altered by air and water, and hence tends to preserve these bodies, just as the bituminized cerecloth preserves an Egyptian mummy; while, on the other hand, the merely woody stem and leaves tend to rot, as fast as the wood of the mummy’s coffin has rotted. Thus the mixed heap of spores, leaves, and stems in the coal-forest would be persistently searched by the long-continued action of air and rain; the leaves and stems would gradually be reduced to little but their carbon, or, in other words, to the condition of mineral charcoal in which we find them; while the spores and sporangia remained as a comparatively unaltered and compact residuum.
There is, indeed, tolerably clear evidence that the coal must, under some circumstances, have been converted into a substance hard enough to be rolled into pebbles, while it yet lay at the surface of the earth; for in some seams of coal, the courses of rivulets, which must have been living water, while the stratum in which their remains are found was still at the surface, have been observed to contain rolled pebbles of the very coal through which the stream has cut its way.