The investigation of the deposits which are now being formed in the basin of the Mediterranean, by the late Professor Edward Forbes, by Professor Williamson and more recently by Dr. Carpenter, and a comparison of the results thus obtained with what is known of the surface fauna, have brought to light the remarkable fact, that while the surface and the shallows abound with Foraminifera and other calcareous shelled organisms, the indications of life become scanty at depths beyond 500 or 600 fathoms, while almost all traces of it disappear at greater depths, and at 1,000 to 2,000 fathoms the bottom is covered with a fine clay.

Dr. Carpenter has discussed the significance of this remarkable fact, and he is disposed to attribute the absence of life at great depths, partly to the absence of any circulation of the water of the Mediterranean at such depths, and partly to the exhaustion of the oxygen of the water by the organic matter contained in the fine clay, which he conceives to be formed by the finest particles of the mud brought down by the rivers which flow into the Mediterranean.

However this may be, the explanation thus offered of the presence of the fine mud, and of the absence of organisms which ordinarily live at the bottom, does not account for the absence of the skeletons of the organisms which undoubtedly abound at the surface of the Mediterranean; and it would seem to have no application to the remarkable fact discovered by the Challenger, that in the open Atlantic and Pacific Oceans, in the midst of the great intermediate zone, and thousands of miles away from the embouchure of any river, the sea-bottom, at depths approaching to and beyond 3,000 fathoms, no longer consists of Globigerina ooze, but of an excessively fine red clay.

Professor Thomson gives the following account of this capital discovery:—­

“According to our present experience, the deposit of Globigerina ooze is limited to water of a certain depth, the extreme limit of the pure characteristic formation being placed at a depth of somewhere about 2,250 fathoms.  Crossing from these shallower regions occupied by the ooze into deeper soundings, we find, universally, that the calcareous formation gradually passes into, and is finally replaced by, an extremely fine pure clay, which occupies, speaking generally, all depths below 2,500 fathoms, and consists almost entirely of a silicate of the red oxide of iron and alumina.  The transition is very slow, and extends over several hundred fathoms of increasing depth; the shells gradually lose their sharpness of outline, and assume a kind of ‘rotten’ look and a brownish colour, and become more and more mixed with a fine amorphous red-brown powder, which increases steadily in proportion until the lime has almost entirely disappeared.  This brown matter is in the finest possible state of subdivision, so fine that when, after sifting it to separate any organisms it might contain, we put it into jars to settle, it remained for days in suspension, giving the water very much the appearance and colour of chocolate.