The Crinoidea, in the abundance of stalked forms in the ancient formations as compared with their present rarity, seem to present us with a fair case of modification from a more embryonic towards a less embryonic condition.  But then, on careful consideration of the facts, the objection arises that the stalk, calyx, and arms of the palaeozoic Crinoid are exceedingly different from the corresponding organs of a larval Comatula; and it might with perfect justice be argued that Actinocrinus and Eucalyptocrinus, for example, depart to the full as widely, in one direction, from the stalked embryo of Comatula, as Comatula itself does in the other.

The Echinidea, again, are frequently quoted as exhibiting a gradual passage from a more generalised to a more specialised type, seeing that the elongated, or oval, Spatangoids appear after the spheroidal Echinoids.  But here it might be argued, on the other hand, that the spheroidal Echinoids, in reality, depart further from the general plan and from the embryonic form than the elongated Spatangoids do; and that the peculiar dental apparatus and the pedicellariae of the former are marks of at least as great differentiation as the petaloid ambulacra and semitae of the latter.

Once more, the prevalence of Macrurous before Brachyurous Podophthalmia is, apparently, a fair piece of evidence in favour of progressive modification in the same order of Crustacea; and yet the case will not stand much sifting, seeing that the Macrurous Podophthalmia depart as far in one direction from the common type of Podophthalmia, or from any embryonic condition of the Brachyura, as the Brachyura do in the other; and that the middle terms between Macrura and Brachyura—­the Anomura—­are little better represented in the older Mesozoic rocks than the Brachyura are.

None of the cases of progressive modification which are cited from among the Invertebrata appear to me to have a foundation less open to criticism than these; and if this be so, no careful reasoner would, I think, be inclined to lay very great stress upon them.  Among the Vertebrata, however, there are a few examples which appear to be far less open to objection.

It is, in fact, true of several groups of Vertebrata which have lived through a considerable range of time, that the endoskeleton (more particularly the spinal column) of the older genera presents a less ossified, and, so far, less differentiated, condition than that of the younger genera.  Thus the Devonian Ganoids, though almost all members of the same sub-order as Polypterus, and presenting numerous important resemblances to the existing genus, which possesses biconclave vertebrae, are, for the most part, wholly devoid of ossified vertebral centra.  The Mesozoic Lepidosteidae, again, have, at most, biconcave vertebrae, while the existing Lepidosteus has Salamandroid, opisthocoelous, vertebrae.  So, none of the Palaeozoic