In the course of my studies I observed that the northern groves, the only ones I was at first acquainted with, were located on just those portions of the general forest soil-belt that were first laid bare toward the close of the glacial period when the ice-sheet began to break up into individual glaciers. And while searching the wide basin of the San Joaquin, and trying to account for the absence of Sequoia where every condition seemed favorable for its growth, it occured to me that this remarkable gap in the Sequoia belt is located exactly in the basin of the vast ancient mer de glace of the San Joaquin and King’s River basins, which poured its frozen floods to the plain, fed by the snows that fell on more than fifty miles of the summit. I then perceived that the next great gap in the belt to the northward, forty miles wide, extending between the Calaveras and Tuolumne groves, occurs in the basin of the great ancient mer de glace of the Tuolumne and Stanislaus basins, and that the smaller gap between the Merced and Mariposa groves occurs in the basin of the smaller glacier of the Merced. The wider the ancient glacier, the wider the corresponding gap in the Sequoia belt.
Finally, pursuing my investigations across the basins of the Kaweah and Tule, I discovered that the Sequoia belt attained its greatest development just where, owing to the topographical peculiarities of the region, the ground had been most perfectly protected from the main ice-rivers that continued to pour past from the summit fountains long after the smaller local glaciers had been melted.
Taking now a general view of the belt, beginning at the south, we see that the majestic ancient glaciers were shed off right and left down the valleys of Kern and King’s rivers by the lofty protective spurs outspread embracingly above the warm Sequoia-filled basins of the Kaweah and Tule. Then, next northward, occurs the wide Sequoia-less channel, or basin, of the ancient San Joaquin and King’s River mer de glace; then the warm, protected spots of Fresno and Mariposa groves; then the Sequoia-less channel of the ancient Merced glacier; next the warm, sheltered ground of the Merced and Tuolumne groves; then the Sequoia-less channel of the grand ancient mer de glace of the Tuolumne and Stanislaus; then the warm old ground of the Calaveras and Stanislaus groves. It appears, therefore, that just where, at a certain period in the history of the Sierra, the glaciers were not, there the Sequoia is, and just where the glaciers were, there the Sequoia is not.
What the other conditions may have been that enabled Sequoia to establish itself upon these oldest and warmest portions of the main glacial soil-belt, I cannot say. I might venture to state, however, in this connection, that since the Sequoia forests present a more and more ancient aspect as they extend southward, I am inclined to think that the species was distributed from the south, while the Sugar Pine,